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Abstract We report a CO(J= 3−2) detection of 23 molecular clouds in the extended ultraviolet (XUV) disk of the spiral galaxy M83 with the Atacama Large Millimeter/submillimeter Array. The observed 1 kpc²region is at about 1.24 times the optical radius (R₂₅) of the disk, where CO(J= 2–1) was previously not detected. The detection and nondetection, as well as the level of star formation (SF) activity in the region, can be explained consistently if the clouds have the mass distribution common among Galactic clouds, such as Orion A—with star-forming dense clumps embedded in thick layers of bulk molecular gas, but in a low-metallicity regime where their outer layers are CO-deficient and CO-dark. The cloud and clump masses, estimated from CO(3−2), range from 8.2 × 10²to 2.3 × 10⁴M_⊙and from 2.7 × 10²to 7.5 × 10³M_⊙, respectively. The most massive clouds appear similar to Orion A in star formation activity as well as in mass, as expected if the cloud mass structure is common. The overall low SF activity in the XUV disk could be due to the relative shortage of gas in the molecular phase. The clouds are distributed like chains up to 600 pc (or longer) in length, suggesting that the trigger of cloud formation is on large scales. The common cloud mass structure also justifies the use of high-JCO transitions to trace the total gas mass of clouds, or galaxies, even in the high-zuniverse. This study is the first demonstration that CO(3−2) is an efficient tracer of molecular clouds even in low-metallicity environments. 

Abstract We present Atacama Large Millimeter/submillimeter Array (ALMA) imaging of molecular gas across the full star-forming disk of the barred spiral galaxy M83 in CO( J = 1–0). We jointly deconvolve the data from ALMA’s 12 m, 7 m, and Total Power arrays using the MIRIAD package. The data have a mass sensitivity and resolution of 10 4 M ⊙ (3 σ ) and 40 pc—sufficient to detect and resolve a typical molecular cloud in the Milky Way with a mass and diameter of 4 × 10 5 M ⊙ and 40 pc, respectively. The full disk coverage shows that the characteristics of molecular gas change radially from the center to outer disk, with the locally measured brightness temperature, velocity dispersion, and integrated intensity (surface density) decreasing outward. The molecular gas distribution shows coherent large-scale structures in the inner part, including the central concentration, offset ridges along the bar, and prominent molecular spiral arms. However, while the arms are still present in the outer disk, they appear less spatially coherent, and even flocculent. Massive filamentary gas concentrations are abundant even in the interarm regions. Building up these structures in the interarm regions would require a very long time (≳100 Myr). Instead, they must have formed within stellar spiral arms and been released into the interarm regions. For such structures to survive through the dynamical processes, the lifetimes of these structures and their constituent molecules and molecular clouds must be long (≳100 Myr). These interarm structures host little or no star formation traced by H α . The new map also shows extended CO emission, which likely represents an ensemble of unresolved molecular clouds. 

Abstract Facultative, heritable endosymbionts are found at intermediate prevalence within most insect species, playing frequent roles in their hosts’ defence against environmental pressures. Focusing onHamiltonella defensa, a common bacterial endosymbiont of aphids, we tested the hypothesis that such pressures impose seasonal balancing selection, shaping a widespread infection polymorphism. In our studied pea aphid (Acyrthosiphon pisum) population,Hamiltonellafrequencies ranged from 23.2% to 68.1% across a six‐month longitudinal survey. Rapid spikes and declines were often consistent across fields, and we estimated that selection coefficients forHamiltonella‐infected aphids changed sign within this field season. Prior laboratory research suggested antiparasitoid defence as the majorHamiltonellabenefit, and costs under parasitoid absence. While a prior field study suggested these forces can sometimes act as counter‐weights in a regime of seasonal balancing selection, our present survey showed no significant relationship between parasitoid wasps andHamiltonellaprevalence. Field cage experiments provided some explanation: parasitoids drove modest ~10% boosts toHamiltonellafrequencies that would be hard to detect under less controlled conditions. They also showed thatHamiltonellawas not always costly under parasitoid exclusion, contradicting another prediction. Instead, our longitudinal survey – and two overwintering studies – showed temperature to be the strongest predictor ofHamiltonellaprevalence. Matching some prior lab discoveries, this suggested that thermally sensitive costs and benefits, unrelated to parasitism, can shapeHamiltonelladynamics. These results add to a growing body of evidence for rapid, seasonal adaptation in multivoltine organisms, suggesting that such adaptation can be mediated through the diverse impacts of heritable bacterial endosymbionts.